My take on this unfortunate event:
Primary cause was improperly torque wheel bolts.
I've crossed that bridge a few times, and it is a long, squirmy, ride.
Because of the depth of the "channels" in the bridge grate, the tires tend to jump from one channel to another, back and forth. That may have facilitated the predisposed loose wheel bolts to jiggle and "walk out" causing the failure to happen on the bridge.
Wikipedia: "Opened in 1957, the 26,372-foot-long (4.995 mi; 8.038 km) [1] bridge is the world's 29th-longest main span and is the longest suspension bridge between anchorages in the Western Hemisphere."